Russian Journal of Theriology. Main page    

Russian Journal of Theriology. Main page
Free access to the published articles
Information about online submission, Articles format, Instructions for authors etc
Instructions for reviewers
Subscription and prices

Русскоязычный вариант сайта
Late Miocene Indarctos punjabiensis atticus (Carnivora, Ursidae) in Ukraine with survey of Indarctos records from the former USSR
Baryshnikov G.F.
P. 83-89
The detailed description of the latest Miocene (MN 13) maxillary fragment with P4-M2 of Indarctos bear from Ananiev in the southern Ukraine is given. The specimen is assigned to the subspecies I. punjabiensis atticus (Weithofer, 1888), comb. nov. A pronounced similarity revealed in tooth proportions and size between I. p. atticus and I. oregonensis from the North American late Miocene suggests I. oregonensis to be only a subspecies: I. punjabiensis oregonensis Merriam, Stock et Moody, 1916, comb. nov. The Indarctos records from the territory of the former USSR are reviewed.

DOI: 10.15298/rusjtheriol.1.2.02


  • Beaumont G. de. 1988. Contributions à l'étude du gisement Miocène supérieur de Montredon (Herault). Les grands mammifères. 2 - Les Carnivores // Palaeovertebrata. Mémoire extraordinaire. P.15-42.
  • Crusafont Pairó M. & Kurtén B. 1976. Bears and bears-dogs from the Vallesian of the Vallés-Penedés Basin, Spain // Acta Zoologica Fennica. No.144. P.1-29.
  • Dames W. 1883. Über das Vorkommen von Hyaenarctos in den Pliocän-Ablagerungen von Pikermi bei Athen // Sitzungsberichte der Gesellschaft naturforschender Freunde zu Berlin. No.8. P.132-139.
  • Dehm R. 1950. Die Raubtiere aus dem Mittel-Miocän (Burdigalium) von Wintershof-West bei Eichstätt in Bayern // Abhandlungen der Bayerischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Klasse. Neue Folge. Hf.58. S.1-141.
  • Dmitrieva E.L. & Nesmeyanov S.A. 1982. [Mammals and stratigraphy of continental Tertiary deposits of the South-West of Middle Asia] // Trudy Paleontologicheskogo Instituta AN SSSR. T.193. P.1-140 [in Russian].
  • Dubrovo I.A. 1962. [Family Ursidae] // Gromova V.I. (ed.). [Fundamentals of Paleontology. Mammals]. Moskva: Gosudarstvennoe Nauchno-Tekhnicheskoe Izdatel'stvo Literatury po Geologii i Okhrane Nedr. P.214-218 [in Russian].
  • Gabunia L.K. 1959. [To history of Hipparion]. Moskva: Izdatel'stvo Akademii Nauk SSSR. 571 p. [in Russian].
  • Ginsburg L. 1999. Order Carnivora // Rössner G. & Heissig K. (eds.). Miocene Land Mammals of Europe. München: Verlag Dr. Friedrich Pfeil. P.109-148.
  • Guérin C. 1980. Les rhinocéros (Mammalia, Perissodactyla) du Miocene terminal au Pleistocene superieur en Europe occidentale: Comparaison avec les espèces actuelles // Documents des Laboratoires de Géologie Lyon. No.79. Fasc.1. P.1-421.
  • Helbing H. 1932. Über einen Indarctos-Schädel aus dem Pontien der Insel Samos. Nebst einem Anhang: Hyaenarctos spec. aus dem Pliocaen von Vialette (Haute-Loire) // Abhandlungen der schweizerischen palaeontologischen Gesellschaft. Bd.52. S.1-18.
  • Hendey Q.B. 1980. Agriotherium (Mammalia, Ursidae) from Langebaanweg, South Africa, and relationships of the genus // Annals of the South African Museum. Vol.81. No.1. P.1-109.
  • Hunt R.M., Jr. 1998. Ursidae // Janis C., Scott K. & Jacobs L. (eds.). Evolution of Tertiary Mammals of North America. Vol.1. Terrestrial Carnivores, Ungulates, and Ungulatelike Mammals. Cambridge University Press. P.174-194.
  • Korotkevich E.L. 1988. [History of formation of Hipparion fauna in Eastern Europe]. Kiev: Naukova Dumka. 164 p. [in Russian].
  • Lungu A.N. & Chemyrtan G.D. 1996. [New species of the genus Indarctos (Carnivora, Mammalia) from the Middle Sarmatian Hipparion fauna of Moldova] // Sokolov V.E. (ed.). [State of the Theriofauna in Russia and Adjacent Territories]. Moskva: Teriologicheskoe Obshchestvo. P.184-190 [in Russian, with English summary].
  • Matthew W.D. 1929. Critical observations upon Siwalik mammals // Bulletin of the American Museum of Natural History. Vol.56. P.437-560.
  • Merriam J.C., Stock Ch. & Moody C. 1916. An American Pliocene bear // University of California Publications. Bulletin of the Department of Geology. Vol.10. No.7. P.87-109.
  • Petter G. & Thomas H. 1986. Les Agriotheriinae (Mammalia, Carnivora) néogènes de l'ancien monde presence du genre Indarctos dans la faune de Menacer (ex-Marceau), Algérie // Geobios. No.19. Fasc.5. P.573-586.
  • Pevzner M.A., Lungu A.N., Vangengeim E.A. & Basilyan A.E. 1987. [Position of Vallesian localities of Hipparion fauna from Moldavia on magnetochronological scale] // Izvestiya AN SSSR. Seriya Geologicheskaya. No.4. P.50-59 [in Russian].
  • Pevzner M.A., Vangengeim E.A., Zhegallo V.I., Zazhigin V.S. & Liskun I.G. 1982. [Correlation of late Neogene beds of Central Asia and Europe on the basis of paleomagnetic and biostratigraphical data] // Izvestiya AN SSSR. Seriya Geologicheskaya. No.6. P.5-16 [in Russian].
  • Pilgrim G.E. 1914. Further description of Indarctos salmontanus Pilgrim, the new genus of bear from the Middle Siwaliks, with some remarks on the fossil Indian Ursidae // Records Geological Survey of India. Vol.44. Pt.3. P.225-233.
  • Qiu Z. & Schmidt-Kittler N. 1983. Agriotherium intermedium (Stach, 1957) from a Pliocene fissure filling of Xiaoxian county (Anhuei province, China) and the phylogenetic position of the genus // Palaeovertebrata. T.13. No.3. P.65-81.
  • Roth C.H. & Morlo M. 1997. Die Raubtiere (Mammalia, Carnivora) aus dem Turolium von Dorn-Dürkheim 1 (Rheinhessen). Teil 2: Ursidae // Courier Forschungs-Institut Senckenberg. Bd.197. S.49-71.
  • Roussiakis S. 2001. Postcranial remains of Indarctos atticus (Ursidae, Mammalia) from the classical locality of Pikermi (Attica, Greece), with a description of the front limb // Senckenbergiana lethaea. Bd.81. S.347-358.
  • Thenius E. 1959. Indarctos arctoides (Carnivora, Mammalia) aus dem Pliozän Österreich nebst einer Revision der Gattung // Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen. Bd.108. Hf.3. S.270-295.
  • Tleuberdina P.A. 1988. [Main localities of Hipparion fauna in Kazakhstan and their biostratigraphical correlation] // Materialy po Istorii Fauny i Flory Kazakhstana. T.10. Alma-Ata. P.38-73 [in Russian].
  • Vereshchagin N.K. 1959. [The Mammals of the Caucasus: A History of the Evolution of the Fauna]. Moskva-Leningrad: Izdatel'stvo Akademii Nauk SSSR. 704 p. [in Russian].
  • Vislobokova I.A. 1990. [Fossil deers of Eurasia] // Trudy Paleontologicheskogo Instituta AN SSSR. T.240. P.1-208 [in Russian with English summary].
  • Vislobokova I.A., Sotnikova M.V. & Dodonov A.E. 2001. Late Miocene - Pliocene mammalian faunas of Russia and neighbouring countries // Bullettino della Società Paleontologica Italiana. Vol.40. No.2. P.307-313.
  • Weithofer A. 1888. Beiträge zur Kenntnis der Fauna von Pikermi bei Athen // Beiträge zur Paläontologie und Geologie Österreich-Ungarn und des Orients. Bd.6. Hf.3. S.225-292.
  • Wolff R.G. 1978. Function and phylogenetic significance of cranial anatomy of an early bear (Indarctos) from Pleistocene sediments of Florida // Carnivore. Vol.1. No.3/4. P.1-12.
  • Zdansky O. 1924. Jungtertiäre Carnivoren Chinas // Palaeontologia Sinica. Series C. Vol.2. Fasc.1. S.1-149.

Download PDF