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Mammalian humerus from the Early Cretaceous of West Siberia.
Gambaryan P.P ., Averianov A.O., Maschenko E.N., Leshchinskiy S.V.
P. 51-58
A distal humerus fragment from the Early Cretaceous Shestakovo locality in Kemerovo Province, West Siberia is described and referred to “Symmetrodonta”. This humerus is characterized by relatively narrow distal epiphysis, large entepicondylar foramen, globular radial condyle, ellipsoidal ulnar condyle, coalesced articular surfaces of both condyles, and a unique additional articular platform for the radius laterally to the radial condyle. This structure of the elbow joint is structurally intermediate between cynodonts, morganucodontids, and multituberculates on the one hand and crown Theria on the other. Among mammals with the similar structure of the elbow joint the humerus from Shestakovo is most similar with the humerus of the Early Cretaceous “symmetrodont” Zhangheotherium from China, and it might belong to a new tinodontid “symmetrodont” being described from the same locality on dental remains.


  • Engelmann G.F. & Callison G.L. 1998. Mammalian faunas of the Morrison Formation // Carpenter K., Chure D.J. & Kirkland J.I. (eds.). The Upper Jurassic Morrison Formation: An Interdisciplinary Study. Modern Geology. Vol.23. P.343-379.
  • Gambaryan P.P. & Kielan-Jaworowska Z. 1997. Sprawling versus parasagittal stance in multituberculate mammals // Acta Palaeontologica Polonica. Vol.42. No.1. P.13-44.
  • Hu Y.-M., Wang Y.-Q., Luo, Z.-X. & Li, C.-K. 1997. A new symmetrodont mammal from China and its implications for mammalian evolution // Nature. Vol.390. P.137-142.
  • Hu Y.-M., Wang Y.-Q., Li C.-K. & Luo Z.-X. 1998. Morphology of dentition and forelimb of Zhangheotherium // Vertebrata PalAsiatica. Vol.36. No.2. P. 102-125.
  • Jenkins F.A. Jr. 1970. Limb movements in a monotreme (Tachyglossus aculeatus): a cineradiographic analysis // Science. Vol.168. P.1473-1475.
  • Jenkins F.A. Jr. & Schaff C.R. 1988. The Early Cretaceous mammal Gobiconodon (Mammalia, Triconodonta) from the Cloverly Formation in Montana // Journal of Vertebrate Paleontology. Vol.8. No.1. P.1-24.
  • Ji Q., Luo Z.-X. & Ji S.-A. 1999. A Chinese triconodont mammal and mosaic evolution of mammalian skeleton // Nature. Vol.398. P.326-330.
  • Kielan-Jaworowska Z. 1998. Humeral torsion in multituberculate mammals // Acta Palaeontologica Polonica. Vol.43. No.1. P.131-134.
  • Kielan-Jaworowska Z. & Gambaryan P.P. 1994. Postcranial anatomy and habits of Asian multituberculate mammals // Fossils and Strata. No.36. P.1-92.
  • Krebs B. 1991. Das Skelett von Henkelotherium guimarotae gen. et sp. nov. (Eupantotheria, Mammalia) aus dem Oberen Jura von Portugal // Berliner geowissenschaftliche Abhandlungen, Reihe A: Geologie und Paläontologie. Bd.133. S.1-121.
  • Lopatin A.V., Masсhenko E.N., Averianov A.O., Rezvyi A.S., Skutschas P.P. & Leshchinskiy S.V. In press. Early Cretaceous mammals from Western Siberia: 1. Tinodontidae // Paleontologicheskii Zhurnal. [in Russian].
  • Martin T. 2000. The dryolestids and the primitive peramurid from the Guimarota Mine // Martin T. & Krebs B. (eds.). Guimarota: A Jurassic Ecosystem. Munich: Verlag Dr. Friedrich Pfeil. P.109-120.
  • Maschenko E.N. 1999a. [First record of an Early Cretaceous triconodont mammal (Mammalia, Triconodonta) in Siberia] // Byuleten' Moskovskogo Obschestva Ispytatelei Prirody, Otdel Geologicheskii. T.24. No.1. P.80 [in Russian].
  • Maschenko E.N. 1999b. [Mesozoic triconodont on the territory of Siberia] // Priroda. No.7. P.52-53 [in Russian].
  • Maschenko E.N. & Lopatin A.V. 1998. First record of an Early Cretaceous triconodont mammal in Siberia // Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, Sciences de la Terre. T.68. P.233-236.
  • Maschenko E.N., Lopatin A.V. & Voronkevich A.V. 2000. [New Early Cretaceous mammals from the locality Shestakovo (Kemerovo Province)] // [Abstracts of Reports of 46th Session of the Paleontological Society]. Sankt Peterburg. P.61-62 [in Russian].
  • Maschenko E.N., Lopatin A.V. & Voronkevich A.V. 2002a. [New Early Cretaceous mammal from Western Siberia] // Doklady Akademii Nauk. T.386. No.5. P.715-717 [in Russian].
  • Maschenko E.N., Lopatin A.V. & Voronkevich A.V. 2003. A new genus of the tegotheriid docodonts (Docodonta, Tegotheriidae) from the Early Cretaceous of West Siberia // Russian Journal of Theriology. Vol.1. No.2. P.75-81.
  • Maschenko E.N. & Voronkevich A.V. 2001. [New data on the diversity of mammals from the Early Cretaceous localities Shestakovo (south-west of Western Siberia)] // [Evolution of Life on the Earth. Materials of the Second International Symposium, November 12-15, 2001]. Tomsk: Izdatel'stvo NTL. P.448-450 [in Russian].
  • Maschenko E.N., Voronkevich A.V., Leshchinskiy S.V. & Fayngerts A.V. 2002b. Early Cretaceous Shestakovo assemblage of mammalian fossils from Western Siberia // First International Palaeontological Congress (IPC2002). Geological Society of Australia, Abstracts. Sydney: McPherson's Printing Group. P.239-241.
  • Pridmore P.A. 1985. Terrestrial locomotion in monotremes (Mammalia: Monotremata) // Journal of Zoology, London. Vol.205. P.53-73.
  • Sereno P.C. & McKenna M.C. 1995. Cretaceous multituberculate skeleton and the early evolution of the mammalian shoulder girdle // Nature. Vol.377. P.144-147.
  • Vázquez-Molinero R., Martin T., Fischer M.S. & Frey R. 2001. Comparative anatomical investigations of the postcranial skeleton of Henkelotherium quimarotae Krebs, 1991 (Eupantotheria, Mammalia) and their implications for its locomotion // Mitteilungen aus dem Museum für Naturkunde in Berlin, Zoologische Reihe. Bd.77. Hf.2. P.207-216.

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