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Pleistocene Mustelidae (Carnivora) from Paleolithic site in Kudaro Caves in the Caucasus
Baryshnikov G.F.
P. 75-95
Kudaro Paleolithic sites in the Southern Ossetia are found to contain five species of mustelids: Meles meles, Martes foina, Martes martes, Vormela peregusna, and Mustela nivalis. All these species occur in the Caucasus at present time; no difference was revealed in the species composition between faunas of Middle and Late Pleistocene. The fossil and recent populations demonstrate a considerable morphological similarity, except weasel represented by a distinct fossil subspecies M. nivalis kudarensis subsp.n.


  • Abramov A.V. 2003. The head colour pattern of the Eurasian badgers (Mustelidae, Meles) // Small Carnivore Conservation. No.29. P.5-7.
  • Abramov A.V. & Baryshnikov G.F. 2000. Geographic variation and intraspecific taxonomy of weasel Mustela nivalis (Carnivora, Mustelidae) // Zoosystematica Rossica, 1999. Vol.8. No.2. P.365-402.
  • Abramov A.V. & Medvedev S.G. 2003. Notes on zoogeography and taxonomy of the badgers (Carnivora: Mustelidae: Meles) and some of their fleas (Siphonaptera: Ceratophyllidae: Paraceras) // Zoosystematica Rossica. T.11. No.2. P.397-402.
  • Abramov A.V. & Pusachenko A.Yu. 2005. Sexual dimorphism of cranial characters in Eurasian badgers, Meles spp. (Carnivora, Mustelidae) // Zoologischer Anzeiger. T.244. P.11-29.
  • Abramov A.V. & Pusachenko A.Yu. 2006. [Geographical variability of skull and taxonomy of Eurasian badgers (Mustelidae, Meles)] // Zoologicheskii Zhurnal. T.85. No.5. P.641-655 [in Russian].
  • Anderson E. 1970. Quaternary evolution of the genus Martes (Carnivora, Mustelidae) // Acta Zoologica Fennica. Vol.130. P.1-132.
  • Argant A. 2004. Les Carnivores du gisement Pliocène final de Saint-Vallier (Drôme, France) // Geobios. T.37. P.133-182.
  • Aristov A.A. & Baryshnikov G.F. 2001. [Mammals of Russia and Adjacent Territories. Carnivores and Pinnepeds]. Sankt-Peterburg. 560 p. [in Russian].
  • Baryshnikov G.F. 1998. Cave bears from the Paleolithic of the Greater Caucasus // Saunders J.J., Styles B.W. & Baryshnikov G.F. (eds.). Quaternary Paleozoology in the Northern Hemisphere. Springfield. P.69-118. (Illinois State Museum Scientific Papers. Vol. 27.)
  • Baryshnikov G. 1999. Bone assemblages from Acheulean and Mousterian levels in the Kudaro Caves in the Caucasus Mountains // The Role of Early Humans in the Accumulation of European Lower and Middle Palaeolithic Bone Assemblages. Monographien des romisch-germanischen Zentralmuseums, Mainz. Bd.42. P.237-253.
  • Baryshnikov G. 2002. Local biochronology of Middle and Late Pleistocene mammals from the Caucasus // Russian Journal of Theriology. Vol.1. No 1. P.61-67.
  • Baryshnikov G.F. & Abramov A.V. 1997. [Structure of baculum (os penis) in Mustelidae (Mammalia, Carnivora). Comunication 1] // Zoologicheskii Zhurnal. T.76. No 9. P.84-97 [in Russian].
  • Baryshnikov G.F. & Golovanova L.V. 1989. [Mammals of the Mousterian site Matuzka in the Kuban Caucasus] // Trudy Zoologicheskogo Instituta Akademii nauk SSSR. Т.198. P.3-55 [in Russian].
  • Baryshnikov G.F. & Potapova O.R. 1990. [Dental variation of badgers (Meles, Carnivora) of the fauna USSR] // Zoologicheskii Zhurnal. Т.69. No.9. P.84-97 [in Russian].
  • Baryshnikov G., Puzachenko A. & Abramov A. 2003. New analysis of variability of check teeth in Eurasian badgers (Carnivora, Mustelidae, Meles) // Russian Journal of Theriology. 2002. Vol.1. No.2. P.133-149.
  • Bonifay M.-F. 1971. Carnivores quaternaries du sud-est de la France // Mémoires du Muséum National d'Histoire Naturelle, ser. C. T.21. Fasc.2. P.43-377 + XXVII pls.
  • Burchak-Abramovich N.I. & Bendukidze O.G. 1971. [Fossil glutton in the Caucasus] // Musei Macedonici Scientiarum Naturalium. T.8. No.12. P.105-111 [in Russian].
  • Gerasimov S. 1985. Species and sex determination of Martes martes and Martes foina by use of systems of craniometrical indices developed by stepwise discriminant analysis // Mammalia. T.49. No.2. P.235-248.
  • Gromova V. 1950. [Determination key to mammals of USSR based on postcranial bones. Part 1. Determination key based on long bones]. Moskva-Leningrad: Izdatel'stvo Akademii Nauk SSSR. 240 p. [in Russian].
  • Guerin C., Barychnikov G. & Mejloumian S. 1992. Survivance tardive d'une forme archaique dans le Pleistocene du Caucase: le Dicerorhinus etruscus brachycephalus du gisement mousterien d'Erivan 1 (Armenie) (Mammalia, Rhinocerotidae) // Bulletin mensuel de la Societé linneenne de Lyon. T.61. Fasc.6. P.166-174.
  • Heptner V.G., Naumov N.P., Yurgenson P.B., Sludsky A.A., Chirkova A.F. & Bannikov A.G. 1967. [Mammals of Soviet Union. T.2. Part 1. Sea Cows and Carnivores]. Moskva: Vysshaya Shkola. 1004 p. [in Russian].
  • Hugueney M. 1975. Les mustelidés (Mammalia, Carnivora) du gisement pléistocene moyen de la Fage (Correze) // Nouvelles Archives du Muséum d'Histoire Naturelle de Lyon. Fasc.13. P.29-46.
  • Knapp M., Rohland N., Weinstock J., Baryshnikov G., Sher A., Doris N., Rabeder G., Pinhasi R., Schmitt H. & Hoffreiter M. 2009. Fist DNA sequences of Asian cave bear fossils reveal deep divergences and complex phylogeographic patterns // Molecular Ecology. Vol.18. No.6. P.1225-1238.
  • Kurose N., Abramov A.V. & Masuda R. 2005. Comparative phylogeography between the ermine Mustela erminea and the least weasel M. nivalis of Palaearctic and Nearctic Regions, based on analysis of mitochondrial DNA control region sequences // Zoological Science. Vol.22. No.10. P.1069-1078.
  • Kurose N., Kaneko Y., Abramov A.V., Siriaroonrat B. & Masuda R. 2001. Low genetic diversity in Japanese populations of the Eurasian badger Meles meles (Mustelidae, Carnivora) revealed by mitochondrial cytochrome b gene sequences // Zoological Science. Vol.18. P.1145-1151.
  • Lioubine V.P. 1998. [The Acheulian epoch of in the Caucasus]. Sankt-Peterburg: Peterburgskoe Vostokovedenie. 192 p. [in Russian].
  • Marmi J., López-Giráldez F., MacDonalds D.W., Calafell F., Zholnerovskaya E., Domingo-Roura X. 2006. Mitochondrial DNA reveals a strong phylogeographic structure in the badger across Eurasia // Molecular Ecology. Vol.15. P.1007-1020.
  • Mashchenko E.N. & Baryshnikov G.F. 2002. The taxonomic status of a macaque (Primates, Cercopithecidae) from the Middle Pleistocene of Georgia // Paleontological Journal. Vol.36. No.4. P.403-413.
  • Nesmeyanov S.A. 1999. [Geomrphological aspects of Paleolithic palaeoecology of the Western Caucasus)]. Moskva: Nauchnyi Mir. 392 p. [in Russian].
  • Opatrný E. 1972. Revise kraniometrický rozdilů na lebkách kuny skalnỉ, Martes foina (Erxleben, 1777), a kuny lesni, Martes martes (Linnaeus, 1758) (Carnivora, Mustelidae) // Acta Universitatis Palackianae Olomucensis. Facultas Rerum Naturalium. Vol.39. P.107-113.
  • Rabeder G. 1976. Die Carnivoren (Mammalia) aus dem Altpleistozän von Deutsch-Altenburg 2. Mit Beiträgen zur Systematik einiger Musteliden und Caniden // Beiträge zur Paläontologie Österreichs, Wien. Bd.1. S.5-119.
  • Sato J.J., Hosoda T., Wolsan M., Tsuchiya K., Yamamoto Y. & Suzuki H. 2003. Phylogenetic relationships and divergence time among mustelids (Mammalia: Carnivora) based on nucleotide sequences of the nuclear interphotoreceptor retinoid binding protein and mitochondrial cytochrome b genes // Zoological Science. Vol.20. P.243-264.
  • Smirnov N.A. 1918. [On the find of glutton mandible during cave excavation in Kutaisi region] // Izvestia Kavkazskogo Museya. T.11. No 3-4. P.1-2 [in Russian].
  • Stone K.D. & Cook J.A. 2002. Molecular evolution of Holarctic martens (genus Martes, Mammalia: Carnivora: Mustelidae) // Molecular Phylogenetics and Evolution. Vol.24. P.169-179.
  • Taberlet P. & Bouvet J. 1994. Mitochondrial DNA polymorphism, phylogeography, and conservation genetics of the brown bear Ursus arctos in Europe // Proceedings of the Royal Society of London, Series B. Vol.255. P.195-220.
  • Vekua A.K. 1978. [Fossil vertebrates from Tsutskhvati caves] // Davitaya F.F. (ed.) [Study of the Caves in Colchis. Tsutskhvatskaya Multilayer Cave System.] Tbilisi: Metsniereba. P.94-126 [in Russian].
  • Vereshchagin N.K. 1959. [Mammals of Caucasus. History of Fauna Origin]. Moskva-Leningrad: Izdatel'stvo Akademii Nauk SSSR. 704 p.
  • Wolsan M., Ruprecht A.L. & Buchalczyk T. 1985. Variation and asymmetry in the dentition of the pine and stone martens (Martes martes and M. foina) from Poland // Acta Theriologica. Vol.30. No 3. P.79-114.

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